Most papers agree that E-M215 and E-M35 originate somewhere in NEA, particularly East Africa which is in sub-Saharan Africa, even E- V38.This is what you wrote
E-M78 & E-Z827 are both sub clades of E-M35. If E-M215 is from the Horn (despite zero evidence of this) why is it both E-M78 (found extremely far west in NW Africa 15k years ago) and E-Z827 found in the Southern Levant 11k years ago? Why does E-V2729 the brother clade to E-M35 seem like a deep Saharan lineage?
E-M215 has nothing to do with SSA. It’s clearly a Saharan ANA lineage. I would even say upstream E- V38 is likely ANA as well and is responsible for the ANA found in Niger-Congo speakers & Mota. The ANA in groups like the Nilotes seems female mediated tho.