Apollo
VIP
I would personally say the Y16897 mutation probably happened up north in S/Western Asia rather that Yemen or Eastern Arabia? It is quite old, and we share it with Europeans as well, so a neolithic dispersal from the Levant/Anatolia would be my first guess.
As to Y44591, I'd say Arabia.
I think that the T-Y28685 group are Ashkenazi Jews (not truly Euros, recent migrations), while the other European ones probably come from Neolithic Anatolians.